Care was taken to form groups balanced in terms of the students' educational background. This encouraged students to help one another.

All students

Bruno Miguel Cardoso Martins


Presently: Post-doctoral research associate at the Sainsbury Laboratory, University of Cambridge
2008-2013: PhD in Systems Biology, University of Edinburgh, supervised by Peter Swain
2007-2013: PhD student at PDBC, Instituto Gulbenkian de Ciência, Oeiras
2006-2007: Junior researcher at Dep. Mechanical Engineering, Instituto Superior Técnico - Technical University of Lisbon

PhD Project

Supervisor: Peter Swain

Cosupervisor: Ramon Grima

Laboratory: Swain Lab - Center for Systems Biology at Edinburgh

University: The University of Edinburgh


Mathematical modelling of signal sensing and transduction: revisiting classical mechanisms. The University of Edinburgh (2013) pp.178.


Bruno M.C. Martins, Peter S. Swain. Ultrasensitivity in Phosphorylation-Dephosphorylation Cycles with Little Substrate. PLoS Comput Biol (2013) 9(8) , e1003175.

Bruno M.C Martins, Peter S Swain. Trade-Offs and Constraints in Allosteric Sensing. PLoS Comput Biol (2011) 7 (11), e1002261.